>>102099575it makes no biological sense at all
but basically, categorizing the whole race from an evolutive perspective:
1. completely uncategorized biology (worms who develop skeletons or exoskeletons based of unknown conditions, with heavily developed brains) divided in different genetic castes:
a. breeding, (intelligent?) immobile grub called mother grub, which undergoes two (known) stages of growth - young virgin grub and mother grub. it seems to reproduce by self
b. fertile, intelligent, fleshy trolls which undergo two stages of growth (grub and humanoid) in charge of essentially terraforming the environment to better fit the needs of the species. these trolls are subdivided into further castes which exhibit different lifespans and, mental abilities and even paranormal powers
c. sterile, non intelligent, exoskeleton possessing drones that ensure the mother grub is protected and ensure it gets inseminated, by force if necessary. probably undergo the same stage of growth the trolls do.
d. several barely mentioned variants of unknown biological function that seem to be genetically tampered forms of drones (ie the judge on the terezi ancestor story)
2. trolls produce what we can assume is a seminal fluid consisting of a pairing half of their genetic code, this code is the mixed with another troll's. by itself the combined fluid doesn't produce an egg, or maybe it produces a dormant egg
3. the drones bring this fluid and inseminate the mother grub with it. the mother grub randomly selects different pairs of fluid for it's new batch of grubs. this fluid doesn't seem to rot or be lost, as descendants of the same troll can be born millennia later
4. the trolls posses to genetic versions of purely sexual traits: male and female. the difference in biology seems to be gaining non-mammary sacks of grease in the chest and general human female anatomy. But it is assumed or implied that their sexual organs are the same.