>>11167199Seems somewhat reasonable. But I have problems with some details. If the main host was an archaeon, then why do eukaryotes have bacterial cell membranes, and not archaeal membranes? The nucleus must come before mitochondria, and the nucleus must be the starter of phagocytosis, meiosis. Bacteria and archaea are not predators by any means. They are chemical transformers, so to speak. They divide into separate cells to avoid predation and viruses. Deep underground, prokaryotes form huge cells with many of rings of DNA, and plasmids are technically no different than the main loop. Thermoreductive evolution seems also reasonable. There is also evidence that big cells of eukaryotic size were present on Earth as early as 3Gya.
According to the viral origin, the nucleus holds hostage the ribosomal ancient genes in a nucleolus, and uses enslaved ribosomes to transcribe nuclear/viral DNA transcripts.